This domain is found in histidyl, glycyl, threonyl and prolyl tRNA synthetases  it is probably the anticodon binding domain .
tRNA synthetases, or tRNA ligases are involved in protein synthesis. This domain is found in histidyl, glycyl, threonyl and prolyl tRNA synthetases [PUBMED:10447505]. It is probably the anticodon binding domain [PUBMED:9115984].
Wolf YI, Aravind L, Grishin NV, Koonin EV; , Genome Res 1999;9:689-710.: Evolution of aminoacyl-tRNA synthetases--analysis of unique domain architectures and phylogenetic trees reveals a complex history of horizontal gene transfer events. PUBMED:10447505 EPMC:10447505 .
Aberg A, Yaremchuk A, Tukalo M, Rasmussen B, Cusack S; , Biochemistry 1997;36:3084-3094.: Crystal structure analysis of the activation of histidine by Thermus thermophilus histidyl-tRNA synthetase. PUBMED:9115984 EPMC:9115984.